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By studying the bones of the smaller toes (as opposed to the big toe), it was observed that their thickness decreased approximately 40,000 to 26,000 years ago.
This led archaeologists to deduce that wearing shoes resulted in less bone growth, resulting in shorter, thinner toes.
The design of shoes has varied enormously through time and from culture to culture, with appearance originally being tied to function.
Additionally, fashion has often dictated many design elements, such as whether shoes have very high heels or flat ones.
Many moccasins were also decorated with various beads and other adornments.
Moccasins were not designed to be waterproof, and in wet weather and warm summer months, most Native Americans went barefoot.
making it the oldest article of clothing discovered in Scandinavia.
It is thought that shoes may have been used long before this, but because the materials used were highly perishable, it is difficult to find evidence of the earliest footwear.
The Romans, who eventually conquered the Greeks and adopted many aspects of their culture, did not adopt the Greek perception of footwear and clothing.
Some shoes are worn as safety equipment, such as steel-soled boots which are required on construction sites.
The world's oldest leather shoe, made from a single piece of cowhide laced with a leather cord along seams at the front and back, was found in the Areni-1 cave complex in Armenia in 2008 and is believed to date to 3500 BC.
The shoe originated in the Catalonian region of Spain as early as the 13th century, and was commonly worn by peasants in the farming communities in the area.
Many medieval shoes were made using the turnshoe method of construction, in which the upper was turned flesh side out, and was lasted onto the sole and joined to the edge by a seam.
The ancient Greeks largely viewed footwear as self-indulgent, unaesthetic and unnecessary.